What factor appears more critical than proximity to the open ocean for inland stopovers?

Answer

Availability of suitable, exposed substrate

Although Whimbrels forage heavily on exposed mudflats and sandy beaches along the coast, their inland stopover behavior indicates a pragmatic focus on resource availability over strict geographical location. When they utilize environments like flooded fields or inland estuaries during migration, the determining factor appears to be substrate condition rather than being right next to the open ocean itself. The critical element enabling them to feed effectively in these inland areas is the availability of suitable, exposed substrate where they can probe for invertebrates, demonstrating that the ability to access buried prey outweighs the need for constant proximity to the vast expanse of the sea.

What factor appears more critical than proximity to the open ocean for inland stopovers?
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