Where does the pupation stage of water beetles typically occur?

Answer

On land, in damp soil near the water's edge

Although water beetles spend most of their lives submerged, their life cycle necessitates a terrestrial phase for transformation. Water beetles undergo complete metamorphosis, and the pupation stage occurs when the larva crawls out of the water. Once out, the larva burrows into the adjacent damp soil or mud bank near the water margin and constructs a protective cell where the transformation into the pupa takes place. This requirement means that nearby, undisturbed banks with suitable substrate are crucial for the successful completion of the reproductive cycle.

Where does the pupation stage of water beetles typically occur?
insectaquaticarthropodwater beetle