What must remain wet for woodlice to breathe using pseudotracheae?
Answer
The pseudotracheae themselves
The pseudotracheae, which are the specialized lung-like respiratory structures derived from pleopods, function by operating fundamentally like gills even when adapted for air. Because of this fundamental biological mechanism, these structures require external moisture to function effectively for gas exchange. If the pseudotracheae dry out, the woodlouse will essentially suffocate, regardless of whether the ambient air temperature is otherwise suitable. This critical dependence on wet respiratory surfaces is the single greatest constraint limiting their successful colonization of drier environments.

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