When did the Oxybelis aeneus clade likely invade South America?

Answer

After the closure of the Panamanian isthmus

The biogeographical distribution of the Oxybelis aeneus clade is linked to major geological events. The clade originated in Central America and subsequently expanded into South America approximately 2.8 million years ago, a dispersal event enabled by the final closure of the Panamanian isthmus, which provided a land bridge between the two continents.

When did the Oxybelis aeneus clade likely invade South America?
taxonomyclassificationsnake