What broad conclusion about location surveying is supported by the difference between rare and common beetles?

For rare beetles, location means precise hydrologic and geologic conditions, while for common beetles, it means general habitat type.

Comparing the habitat needs of highly specialized, rare species versus generalist families reveals a crucial difference in how 'location' must be defined during assessment. Common diving beetle families might be found in nearly any pond within a region, meaning their location is adequately described by a general habitat type designation. In contrast, rare species like *B. hungerfordi* have such stringent requirements—specific flow rates, substrate composition, and lack of siltation—that their location is defined by a few kilometers of perfect lotic conditions. For these rare species, the location description must be highly granular, detailing the precise combination of physical and chemical parameters, or hydrologic and geologic conditions, that define their narrow ecological window.