Which species are White-crowned Sparrows frequently observed associating with in mixed-species winter flocks?

Answer

Juncos, other sparrows, and towhees.

During the non-breeding season, White-crowned Sparrows are known to be social birds that readily join foraging groups composed of other seed-eating species. They frequently associate with ground-foraging birds that share similar habitat preferences and dietary needs. Specifically, it is common to find them mingling with Dark-eyed Juncos, various other species of sparrows, and towhees while they are actively kicking up debris in search of food sources beneath vegetation.

Which species are White-crowned Sparrows frequently observed associating with in mixed-species winter flocks?
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