What role did Dalmatians assume that cemented their iconic niche in American culture after transportation moved away from horses?
Answer
Running ahead of horse-drawn fire engines to act as living sirens
When horse-drawn fire engines were the norm, Dalmatians ran ahead of the noisy apparatus to clear pathways for the horses and men rushing to a blaze, effectively acting as a living siren.

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Dalmatian – Full History - YouTube
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