How do penguin flippers generate underwater propulsion compared to flying birds?

Answer

Propulsion is achieved from both the downstroke and the upstroke.

Unlike flying birds where propulsion is primarily generated during the downstroke, penguin flippers are modified so powerfully that they generate thrust during both the downward sweep and the upward recovery sweep through the water. This requires immense power supplied by their strong chest and back muscles. The skeletal structure reflects this need, featuring flattened, fused, and dense bones that lack the necessary air pockets for aerial lift, making the flippers specialized for continuous underwater movement rather than flight.

How do penguin flippers generate underwater propulsion compared to flying birds?
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