When differentiating between visually similar species like the Macaroni and Royal penguins, what structural aspect of the crests is often the key differentiator?

Answer

The exact attachment point and density of the feathers.

Observing the exact attachment point and density of the crest feathers, rather than just color, can often be the key differentiator between similar species like the Macaroni and Royal penguins.

When differentiating between visually similar species like the Macaroni and Royal penguins, what structural aspect of the crests is often the key differentiator?
birdanimalgenuspenguinEudyptes