What ancestral trait causes an under-exercised Welsh Springer to become frustrated?

Answer

High energy level and desire to be active participants

The high drive and physical capacity required to work tirelessly for hours covering vast ground in the Welsh countryside were bred directly into the modern dog. This necessity for sustained activity has translated into a defining characteristic of the contemporary Welsh Springer: an extremely high energy level coupled with a profound desire to be involved in all family activities. When this inherent requirement for mental and physical stimulation—whether through agility, obedience, or fieldwork—is not met, the dog often becomes frustrated, which can manifest as destructive behaviors.

What ancestral trait causes an under-exercised Welsh Springer to become frustrated?
evolutionDogbreedspanielWelsh