What specific health issue can result from excessive amounts of fruit or animal protein in a tortoise's diet?

Answer

Pyramiding of the shell

Consuming too much fruit or animal protein can lead to severe health problems for tortoises, notably causing the shell structure to become misshapen, known as pyramiding.

What specific health issue can result from excessive amounts of fruit or animal protein in a tortoise's diet?
Shellreptileherbivoretortoiseslow